AP Biology (An Introduction)
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Transcript AP Biology (An Introduction)
Chapter 24
Species & Speciation
Chapter 24 in a nutshell
Microevolution v. Macroevolution
Speciation
Prezygotic
Postzygotic
Gradualism v. Punctuated equilibrium
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Species
Morphological species concept
Divisions of organisms into discrete units called species
If two organisms look different enough, then they are
considered different species
Biological species concept
Population or group of populations whose members have
the potential to interbreed in nature and produce viable,
fertile offspring
This population is unable to produce viable, fertile
offspring with members of another population
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Micro- vs. Macro- Evolutions
Microevolution
Change in the genetic makeup of a population over
generations
Changes (adaptations) confined to a particular gene pool
or population
Macroevolution
Changes (evolution) above the species level
Changes used to define higher taxa
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Speciation
The process that produces a new species
May be the result of microevolution OR
macroevolution
How could microevolution lead to speciation?
How could a new species arise from macroevolution?
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Reproductive Isolation
Biological barriers that impede members of 2 species
from producing viable, fertile offspring
There are 2 types of barriers that lead to reproductive
isolation
Prezygotic
Pre-mating
Post-mating
Postzygotic
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Prezygotic Barriers
Pre-Mating
Habitat isolation
Temporal isolation
Behavioral isolation
Post-Mating
Mechanical isolation
Gametic isolation
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Prezygotic Barriers (Page 1)
Pre-Mating
Habitat isolation
2 species occupy different habitats in the geographic area
Tree-dwelling species, Bush-dwelling species
Temporal isolation
Temporal = time
Species breed at different times
Different time of day, different seasons, or different years
Behavioral isolation
Certain signals or types of behavior are integral to
courtship, and these signals/behaviors are unique to the
species
Other species do not respond to these signals or behaviors
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Boobies!!
(blue-footed)
Different mating season –
late winter or late summer
Different habitats –
water or terrestrial
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Prezygotic Barriers (Post-Mating)
Mechanical isolation
Anatomical incompatibility
Insect copulatory organs are elaborated for a particular
species
Gametic isolation
Even if gametes meet, they are unable to fuse
Sperm may not be able to survive in the reproductive
tract of females from another species
Purple + Red sea urchins cannot mate b/c gametes
cannot fuse
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Postzygotic Barriers
Reduced hybrid vitality
Reduced hybrid fertility
Hybrid Breakdown
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Postzygotic Barriers
Reduced Hybrid vitality
Although zygote forms, genetic incompatibility causes
developmental cessation
In some salamanders, hybrids form but do not complete
development
Reduced Hybrid fertility
Sterile hybrids
Donkey + Horse = mule (Sterile)
Hybrid breakdown
Although first-generation hybrids survive & reproduce,
their offspring are feeble or sterile
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Geographic Barrier
No Geographic Barrier
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Allopatric Speciation
Causes
Geologic events or processes that fragment the
population
Emergence of a mountain range
Formation of a land bridge
Evaporation of a large lake into multiple smaller lakes
Leads to a significant alteration of a gene pool
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Sympatric Speciation
Sympatric = sym (same) + patr (fatherland)
Same country
Geographically overlapping populations
Major mechanism of sympatric speciation is Polyploidy
Autopolyploidy
Allopolyploidy
Preferential Habitation
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Polyploidy
Extra set(s) of chromosomes
Autopolyploidy
> 2n
Typically, nondisjunction (Meiosis) in Plants
All chromosomes from the same species
Example
Tetraploid (4n) + diploid (2n)
Produce triploid (3n) offspring which are sterile
But plant tetraploids can self fertilize or mate with other
tetraploids = viable, fertile polyploid offspring
Tetraploids are reproductively isolated from the diploid
population
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Polyploidy (Page 2)
Allopolyploidy
Plants
2 different species interbreed = hybrid
Hybrids cannot breed with either original species
But hybrids can successfully mate with each other
Self-pollination successfully occurs as well
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Polyploidy (Page 3)
Can happen in animals but not common
In animals, sympatric speciation is usually due to part
of the population switching to a new habitat, food
source, or other resource
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Adaptive Radiation
Many new species arise from a common ancestor given
new environmental opportunities or challenges
Think Darwin’s finches & Galapagos islands
As the finches moved from South America to the
Galapagos, different environments and food sources
were encountered
Hawaiian Plants
As the plants spread to Hawaiian islands, different
environments on different islands = evolution into
several distinct forms
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Hawaiian Plants that had a common
ancestor
Arose from Adaptive Radiation
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Gradualism vs. Punctuated Equilibrium
Gradualism – species descended from a common
ancestor and gradually diverge in morphology
(acquiring unique adaptations) until they are
reproductively isolated
Punctuated Equilibrium – long periods of apparent
stasis punctuated by brief periods of sudden and rapid
change
Fossil record confirms the punctuated equilibrium model
Also eliminates the necessity of finding “missing links”
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