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Invertebrates
BIOZ 341
Dr, Rafat Khattab
Biology Department
E-mail: [email protected]
Invertebrate Zoology
Study of invertebrate animals
Inverts make up at least 99% of all extant (living)
animal species on the planet
Over 1,000,000 described spp. (species) on the
planet (mostly insects)
Estimated 10 to 30 million spp. have yet to be
described
We will cover 19 (time permitting) of the approx. 35
animal phyla
Invertebrates
Invertebrates are those animals without
backbone.
They can be classified into several taxa
according to several criteria:
1- Presence or absence of proper tissues,
2- Number of layers of tissues present
3- Presence or absence of body cavity.
Invertebrates include protists , parazoans
and metazoans orgamnisms
Protista; which includes Protozoa and
other single celled organisms
Parazoa are those multicellular animals,
which lack proper tissues. Animals belong
to Phylum Porifera (sponges) are good
representatives to Parazoa.
Metazoa are those multicellular animals,
which have proper tissues.
Phylum Coelentrata (Cnidaria) includes
diploblastic (two layered) metazoans.
Acoelomates are triploblastic (three
layered) metazoans without a body cavity
(solid bodied invertebrates), such as
Phylum Platyhelminthes.
Pseudocoelomates are triploblastic
metazoans with a pseudocoelom (the
cavity originated from the blastocoel),
such as Phylum Aschelminthes.
Coelomates are triploblastic metazoans
with true coelom (the cavity originated
from the mesoderm).
These include annelids (segmented
worms), arthropods (jointed legs
invertebrates), molluscs (soft bodied
invertebrates), echinoderms (spined skin
invertebrates), and chordates (animals
with a notochord).
This course deals with the study of
taxonomy, morphology, and biology of
some major examples representing
Protozoa, Parazoa, lower and some higher
Metazoa of invertebrates.
Exploring invertebrate diversity
PORIFERA (5,500 species)
A sponge
PLACOZOA (1 species)
CNIDARIA (10,000 species)
A jelly
KINORHYNCHA (150 species)
0.5 mm
250 µm
A placozoan (LM)
PLATYHELMINTHES (20,000 species)
A marine flatworm
ECTOPROCTA (4,500 species)
Figure 33.3
Ectoprocts
A kinorhynch (LM)
ROTIFERA (1,800 species)
A rotifer (LM)
PHORONIDA (20 species)
Phoronids
Exploring invertebrate diversity
BRACHIOPODA (335 species)
A brachiopod
ACANTHOCEPHALA (1,100 species)
NEMERTEA (900 species)
A ribbon worm
CTENOPHORA (100 species)
5 mm
An acanthocephalan
MOLLUSCA (93,000 species)
A ctenophore, or comb jelly
ANNELIDA (16,500 species)
An octopus
LORICIFERA (10 species)
A marine annelid
PRIAPULA (16 species)
50 µm
Figure 33.3
A loriciferan (LM)
A priapulan
Exploring invertebrate diversity
ARTHROPODA (1,000,000 + species)
NEMATODA (25,000 species)
A roundworm
A scorpion (an arachnid)
CYCLIOPHORA (1 species)
TARDIGRADA (800 species)
100 µm
100 µm
A cycliophoran (colorized SEM)
Tardigrades (colorized SEM)
HEMICHORDATA (85 species)
ONYCHOPHORA (110 species)
An onychophoran
An acorn worm
ECHINODERMATA (7,000 species)
Figure 33.3
A sea urchin
CHORDATA (52,000 species)
A tunicate
Preview of Phyla We Will Cover
Protozoa – animal-like
protists
Do not belong to
kingdom animalia, and
thus are not considered
to be invertebrate
animals
Important evolutionary
link between
prokaryotes, and
everyday plants and
Volvox
Phylum Porifera
Sponges
Phylum Cnidaria
Jellyfish, Anemones, and Corals
Portuguese Man O’ War
Phylum Ctenophora
Comb Jellies
Phylum Platyhelminthes
Flatworms
Phylum Nemertea
Ribbon Worms
Phylum Mollusca
Chitons, Clams, Snails, Slugs, Squids, and
Octopi
Banana Slug
Phylum Annelida
Segmented Worms
Phylum Echiura
Spoonworms
Phylum Sipuncula
Peanut Worms
Phylum Tardigrada
Water Bears
Phylum Arthropoda
Horseshoe Crabs, Arachnids, Crustaceans,
Myriapods, and Insects
Phylum Gastrotricha
Name means “stomach hair”
Phylum Nematoda
Roundworms
Caenorhabditis elegans = good
Ascaris lumbricoides = bad
Phylum Rotifera
Wheel bearers
Phylum Phoronida
A lophophorate
Phylum Brachiopoda
Lamp shells, another lophophorate
Phylum Bryozoa
Bryozoans, the largest phylum in the
superphylum Lophophorata
Phylum Echinodermata
Starfish, Brittle Stars, Urchins, and Sea
Cucumbers
Phylum Chordata
Phylum Chordata is
the only phylum
containing
vertebrate animals,
however, there are
some invertebrate
chordates