Transcript Botany-Fern

Chapter 21
Lecture Outline
The Seedless
Vascular Plants:
Ferns and Their
Relatives
Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display.
Outline

Introduction– Primitive Vascular Plants

Phylum Psilotophyta – The Whisk Ferns


Phylum Lycophyta – The Ground Pines,
Spike Mosses and Quillworts
Phylum Equisetophyta – The Horsetails and
Scouring Rushes

Phylum Polypodiophyta – The Ferns

Fossils
Introduction to Primitive Vascular Plants

Primitive Vascular plant characteristics:
• Internal conducting tissue developed.
• True leaves
• Roots
• Gametophytes are smaller .
• Sporophytes are larger and dominant
• Primitive: no seeds

Four phyla of seedless vascular plants:
Psilotophyta, Lycophyta, Equisetophyta,
Polypodiophyta
Introduction

Psilotophyta
• Sporophytes have neither
true leaves, nor roots.
• Stems and rhizomes fork
evenly.
Psilotum

Lycophyta
• Plants covered with
microphylls.
–
Microphylls - Leaves with
single vein whose trace is not
associated with a leaf gap
Lycopodium
Introduction

Equisetophyta
• Sporophytes have ribbed
stems containing silica.
• Have whorled, scalelike
microphylls that lack
chlorophyll

Equisetum
Polypodiophyta
• Sporophytes have
megaphylls that are often
large and much divided.
–
Megaphylls - Leaves with
more than one vein and a leaf
trace associated with leaf gap
A fern
Phylum Equisetophyta – The Horsetails and
Scouring Rushes


Equisetum
SPOROPHYTE:
• Branched and unbranched
forms, usually less than 1.3
meters tall
• Stems jointed and ribbed.
• Green and do
Photosynthesis
• If branched, then branches
in whorls.
• Long-lived
Branched Form

Silica deposits on walls of stem epidermal cells.
Un-branched Form
• Stomata in grooves
between ribs
• Scalelike leaves in
whorls at nodes =
microphylls; are
nonfunctional
• Strobili = Non-woody
Cones

GAMETOPHYTES
• Gametophytes lobed, green, cushionlike, up to 8
mm in diameter.
• Short-lived
• Small
• Have Archegonia &
Antheridia
- make eggs and sperm
Phylum Equisetophyta

Equisetum reproduction:
• Asexual by fragmentation
of rhizomes
• Sexual reproduction:
–
Strobili at tips of stems with
sporangia produce spores.
–
Spores green with 4 ribbonlike elaters attached.
o
Aid in spore dispersal
Spores with
elaters
-- Spores released into ____________
and grow into _______
-- Gametophytes make_____
-- Fertilization = ___________________________and
produces _________
Phylum Equisetophyta

Equisetum reproduction:
Phylum Equisetophyta

Human and ecological
relevance:

STUDENTS DO
Reconstruction of fossil
giant horsetail, Calamites
Phylum Polypodiophyta – The Ferns

Sporophyte Structure and form:
• Vary in size from tiny floating forms less than 1
cm to giant tropical tree ferns up to 25 m tall
–
Fern leaves are megaphylls - Referred to as fronds.
o
Typically divided into smaller segments
Phylum Polypodiophyta – The Ferns
• Sporophyte is
conspicuous phase.
–
Fronds, rhizomes, roots
–
Fronds first appear
coiled in crozier
(fiddlehead), and then
unroll and expand.
o
Fronds often divided
into segments called
pinnae (singular:
pinna).
Crozier
Phylum Polypodiophyta – The Ferns

Reproduction:
• Sporangia stalked.
–
May be scattered on lower
leaf surface, confined to
margins, or found in
discrete clusters called sori
(singular: sorus).
o
–
Sori may be protected by
indusia (singular: indusium).
With row of heavy-walled,
Sorus covered
by indusium
brownish cells = annulus
o Annulus catapults spores out of sporangium
Phylum Polypodiophyta – The Ferns
• Meiosis forms spores in sporangia.
GAMETOPHYTES
• Spores released and grow into gametophytes called
prothalli (singular: prothallus).
• Prothalli are one cell thick,
and have archegonia and
antheridia.
Archegonia
Antheridia
• Require external water for reproduction
because: ____
• Zygote develops into young sporophyte.
• Gametophyte usually dies and leaves sporophyte
growing independently.
Phylum Polypodiophyta – The Ferns

Reproduction:
Phylum Polypodiophyta

Human and ecological relevance:
• House plants
–
Function well as air filters
• Outdoor ornamentals
• Cooked rhizomes as food
• Folk medicine
• Fronds used in thatching for houses.
• Basketry and weaving
Fossils

A fossil - Any recognizable prehistoric
organic object preserved from past
geological ages.
• Conditions of formation almost always include
quick burial and an accumulation of sediments.
–
Hard parts more likely preserved than soft parts.
Review

Introduction

Phylum Psilotophyta – The Whisk Ferns


Phylum Lycophyta – The Ground Pines,
Spike Mosses and Quillworts
Phylum Equisetophyta – The Horsetails and
Scouring Rushes

Phylum Polypodiophyta – The Ferns

Fossils