Tree of Life 1

download report

Transcript Tree of Life 1

Tree of Life
 Planet
Earth is about 4.6 billion years old.
 Oldest known rocks are about 3.8 billion
years old.
 Oldest fossils (prokaryotes) are about 3.5
billion years old.
Tree of Life
 All
living organisms on this planet share a
common ancestor.
The tree of life reflects the branching
pattern of speciation (phylogenetic history
of life) that has occurred since the origin of
Tree of Life
 There
is an excellent Tree of Life website
in which you can trace the branching
pattern of the history of life and explore
Tree of Life
 There
is a hierarchichal classification of
life in which organisms are progressively
nested within larger and larger categories
as more distant relatives are included in
the classification (as explored previously).
 The highest level of classification is the
Domain of which there are three.
Domains Bacteria and Archaea
 Domain
 Domain Archaea
 The
domains Bacteria and Archaea are
both prokaryotes (they have no nucleus
and the DNA is not arranged in
chromosomes). Prokaryote derived from
the Greek Pro meaning before and karyon
meaning a kernel [i.e. a nucleus]
Domain Bacteria
 Includes
most of the bacteria people are
familiar with including disease-causing
species (Salmonella; Vibrio cholerae
which causes cholera), nitrogen-fixing
(Nitrosomonas) and parasites (Borrelia
burgdorferi which causes Lyme disease).
Domain Bacteria
 Bacteria
play a major role in
decomposition and many live symbiotically
with other organisms including humans
helping to break down or synthesize foods
needed by the host.
Domain Archaea
The Archaea include many extremophiles,
organisms that live in extreme environments.
Includes thermophiles which tolerate extreme
heat (e.g. live in geysers and hot springs where
temps may reach 90 degrees celsius) and
halophiles (salt lovers, which live in very saline
environments (e.g. Great Salt Lake, Dead Sea)
Archaea in hot springs
R. Shand, N. Arizona Univ.
Archaea in dead sea
Archaea in hydrothermal vent
Ruth Blake / Yale
Bacteria and Archaea
 Bacteria
and Archaea are both
prokaryotes and their DNA is arranged in
circular structures called plasmids.
 However, they have substantial
differences in their biochemistry, cell wall
structure and other molecular details.
Bacteria vs. Archaea
Bacteria are inhibited by antibiotics Streptomycin
and Chloramphenicol but Archaea are not.
Archaea in common with Eukarya have histone
proteins associated with their DNA, have introns
in their DNA, and have several kinds of RNA
polymerase. Bacteria lack these features.
 Archaea and Eukarya thus are members of a
Domain Eukarya
 Domain
Eukarya contains the eukaryotic
organisms (from Greek eu true and karyon
a kernal) which have a true nucleus and
DNA arranged in chromosomes.
 Eukaryotic
cells are much larger and
complex than prokaryotic cells and contain
organelles such as mitochondria,
chloroplasts, and lysosomes.
Domain Eukarya
 Domain
Eukarya includes three kingdoms
the Plantae, Fungi and Animalia.
 There
are also a number of unicellular
eukaryotes that may form as many as five
other kingdoms. These were formerly
grouped in the Protista.
Domain Eukarya
 Plantae,
Fungi and Animalia are mostly
multicellular, but plants are autotrophic
(produce their own food by
photosynthesis) whereas the fungi and
animals are heterotrophic (consume other
 Fungi
are heterotrophs and feed by
 They
secrete enzymes outside their
bodies (exoenzymes) which break down
complex molecules to simpler ones which
the fungus can absorb.
 Some
fungi are unicellular (yeasts), but
most are multicellular.
 Body
of multicellular fungi made up of tiny
filaments called hyphae.
 The
hyphae form a mass called a
mycelium that penetrates the medium the
fungus is feeding on.
 Mushrooms
and toadstools are the familiar
reproductive structures of fungi.
 Fungi produce spores which may be
sexually or asexually produced
 Fungi
and Animalia share a more recent
common ancestor (about 1.5 billion years
ago) than they do with Plantae.
 Fungi
are believed to have evolved from
flagellated single-celled protistans, which
suggests multicellularity arose
independently in Fungi and Animalia
Ancestor to land plants: Green Algae
Land Plants (Embryophytes):
 Bryophytes
(mosses, etc.)
 Ferns and relatives
 Gymnosperms
 Angiosperms
(Mosses, etc.)
Ferns and fern allies
Plant structure and function
(parts of chapters 35, 36 and 37)
 Unlike
animals, plants remain in one place
and produce food through photosynthesis.
 To
carry out photosynthesis plants must
obtain water and minerals from the soil,
CO2 from the air, and light from the sun.
 The
structure of plants reflects their need
to carry out these tasks.
Basic structure of plants
 Plants
have three basic organs:
 These
organs are organized into two
systems: the largely below-ground root
system and the above-ground shoot
system (stems and leaves).
 Roots
perform several tasks. They
Anchor the plant in place
Absorb minerals and water
Store organic nutrients such as sugars (e.g.
carrot, sugar beet, turnip).
 Roots
systems may have a central
taproot with lateral roots branching off
from it (e.g. dandelion).
 Alternatively,
a root system may have no
obvious main root, but instead be a
fibrous system with many small roots
growing from the stem, each of which has
its own lateral roots (e.g. grasses).
 The
entire root system anchors a plant in
place, but absorption of water and
minerals occurs mainly at the root tips.
 At
the root tips huge numbers of root
hairs increase the surface area
Root hairs
 Root
hairs are extensions of individual
epidermal root cells and are not
multicellular structures
(as lateral roots are).
 Root
hairs are permeable to water and
adhere closely to soil particles allowing
efficient absorption of water and nutrients.
 Most
plants forms mutually beneficial
relationships with fungi, which facilitate
absorption of water and minerals.
The plants and fungi form mycorrhizae:
symbiotic associations of plant roots united with
fungal hyphae (hyphae are tiny filaments that
form the bulk of a fungus).
Most plants form these symbiotic mycorrhizal
relationships and they greatly enhance the
plants growth. [a symbiotic relationship is a
close, mutually beneficial relationship]
Mycorrhizae (white) growing on a root
 The
fungal hyphae grow over the root and
penetrate into it and may in some cases
form a mantle or layer over the root.
 The
fungus benefits from a steady supply
of sugar donated by the host plant.
Plant receives numerous benefits:
Fungus greatly increases surface area for absorption
(can be as much as 3 meters of hyphae per cm of
plant root length).
Fungus selectively absorbs phosphate and other
nutrients and supplies them to plant.
Fungus may secrete growth factors that promote root
Fungus may produce antibiotics that protect the plant
from pathogenic bacteria and fungi in the soil.
 Plant-fungus
symbiosis may have been
one of the early adaptations that allowed
plants to colonize the land, which probably
initially was quite nutrient poor.
 Fossils
of some of the earliest plants show
Shoot Systems
 Shoot
systems consist of stems and
 Stems
are elongated structures comprised
of nodes and internodes.
 Nodes
are where leaves are attached and
internodes are the sections in between.
Shoot Systems
 Stems
have a terminal bud at the tip and
this is where elongation takes place,
enabling the stem to reach upwards
towards the light.
 If
the tip of the stem is eaten or shaded,
however, axillary buds (buds on the side)
will begin to grow.
Shoot Systems
 Gardeners
 By
shape plants by pruning them.
removing terminal buds a bushier plant
can be produced or by removing lateral
flower buds a single large flower can be
Shoot Systems
 Stems
have been greatly modified in many
plants to perform a variety of functions.
 Rhizomes,
bulbs, tubers, and stolons are
all modified stems although they are often
mistaken for roots.
Modified stems
Bulbs: vertical shoots that grow underground.
The “flesh” of a bulb (e.g. an onion) consists of
leaves modified for food storage.
Stolons and rhizomes: are stems that grow on
(stolons) or just under (rhizomes) the soil
surface. New plantlets form periodically along
the length of these stems (asexual
Modified stems
 Tubers:
are enlarged ends of rhizomes
specialized for storing food (e.g. potato).
 The
“eyes” of a tuber are axillary buds.
 Leaves
are the main photosynthetic organ
of plants, although green stems also
perform photosynthesis.
 Leaves
vary in form, but usually have a flat
blade and a stalk (petiole) that joins the
leaf to the stem.
Unlike animals, plants remain in one place
and produce food through photosynthesis.
In the process of photosynthesis plants (and
other photosynthetic organisms such as
algae, other protists, and cyanobacteria) trap
the energy in sunlight and store it in
chemical bonds.
The energy stored in chemical bonds can
then be used to fuel metabolic processes.
Figure 10.2
Plants and photosynthesis
 This
 In
process is called photosynthesis.
this class we will not discuss the
process of photosynthesis in detail. It
is covered in depth in Bio 101.
 In
photosynthesis carbon dioxide (CO2)
and water (H20) and the energy
provided by light are used to make
CO2 + 12 H20 + energy 
C6H12O6 + 6O2 + 6 H20
 The
organelle plants use to carry out
photosynthesis is the chloroplast.
 In
plants chloroplasts are concentrated
in the leaves, which generally are thin
and flat to allow maximum exposure to
Fig 10.3
 Leaves
are generally flat to maximize the
area exposed to the sun and minimize the
distance gases must be transported to and
from photosynthesizing cells.
 However,
in many cases leaves have been
substantially modified by natural selection
to perform other functions.
Modified leaves
Tendrils of climbing plants such as clematis are often
modified leaves.
Spines of cacti are modified leaves (most
photosynthesis being carried out by the fleshy stem.
Some leaves are modifed as storage leaves to store
Some leaves called bracts look like petals (e.g. in
dogwoods) being brightly colored and enlarged to attract
pollinators to the flowers they surround.
Some leaves produce plantlets that drop off the plant
and take root in the soil.
Plant vascular system
 Plants
contain two vascular systems that
transport water, minerals, and sugars
around the plant.
 Xylem
transports water and dissolved
minerals from the roots into the shoots.
 Phloem
transports sugars from the leaves
to where they are needed in the plant.
Plant vascular system
 Xylem
cells are dead at functional maturity
and form thin elongated tubes that water
moves through.
 Phloem
cells are alive.
Plant secondary growth
Primary growth is stem elongation, secondary
growth refers to the thickening of woody plants
over time.
Xylem and phloem cells are both produced by a
plant tissue called vascular cambium that is
located under the bark.
This cambium produces xylem cells on the
inside and phloem on its outside.
Plant secondary growth
 As
the plant grows older inner xylem
tissue forms the heartwood of the tree.
This tissue no longer transports liquid.
 Xylem
cells have thick lignified walls (lignin
is a complex cross-linked polymer) that
provide structural support for the plant.
 The outer (more recently produced) xylem
is called sapwood and this carries liquid.
Plant secondary growth
 On
the outside of the cambium layer
phloem is produced. Phloem is produced
more slowly than xylem and older phloem
is sloughed off the tree so it does not
accumulate as xylem does.
Plant secondary growth
 Because
the outer layer of phloem is
essential to transportation, a tree that is
“ringed” by grazers (i.e., has its outer bark
removed around the circumference of the
plant) will die.
 In
contrast, a tree may be hollowed out
and still survive.