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Community Structure
and Biodiversity
Chapter 40
Ecosystems
Ecosystems vary in size.
They
can be as small as a puddle or as
large as the Earth itself.
Any group of living and nonliving things
interacting with each other can be
considered as an ecosystem.
Within each ecosystem, there are
habitats
These
may also vary in size.
A habitat is the place where a population
lives. - Turlock habitat
A population is a group of living organisms
of the same kind living in the same place at
the same time. Turlock humans
All the different populations interact and
form a community. City of Turlock
The community of living things interacts
with the non-living world around it to
form the ecosystem.
The habitat must supply the needs of
organisms, such as food, water,
temperature, oxygen, and minerals.
If the population's needs are not met, it
will move to a better habitat.
Two different populations can not
occupy the same niche at the same
time, however.
Community
All the populations that live together in
a habitat
Habitat is the type of place where
individuals of a species typically live
Type of habitat shapes a community’s
structure
Factors Shaping
Community Structure
Climate and topography
Available foods and resources
Adaptations of species in community
Species interactions
Arrival and disappearance of species
Physical disturbances
Niche
Sum of activities and relationships in
which a species engages to secure and
use resources necessary for survival
and reproduction
Realized &
Fundamental Niches - What???
Fundamental niche
Theoretical
niche occupied in the absence
of any competing species
Realized niche
Niche
a species actually occupies
Realized niche is some fraction of the
fundamental niche
Species Interactions
Most interactions are neutral; have no effect
on either species
Commensalism helps one species and has
no effect on the other
Mutualism helps both species
Species Interactions
Interspecific competition has a negative
effect on both species
Predation and parasitism both benefit
one species at a cost to another
Symbiosis
Living together for at least some part of
the life cycle
Commensalism, mutualism, and
parasitism are forms of symbiosis
Mutualism
Both species benefit
Many examples in nature
Some mutualisms are obligatory;
partners depend upon each other
Example: Mycorrhizae
Obligatory mutualism between fungus
and plant root
Fungus supplies mineral ions to root
Root supplies sugars to fungus
Example: Yucca and Yucca Moth
Example of an obligatory mutualism
Each species of yucca is pollinated only
by one species of moth
Moth larvae can grow only in that one
species of yucca
Competition
Interspecific - between species
Intraspecific - between members of the
same species
Intraspecific competition is most intense
Forms of Competition
Competitors may have equal access to
a resource; compete to exploit resource
more effectively
One competitor may be able to control
access to a resource, to exclude others
Competitive Exclusion Principle
When two species compete for identical
resources, one will be more successful
and will eventually eliminate the other
Competitive Exclusion Expt.
Protists - living together
Paramecium caudatum
Paramecium aurelia
Resource Partitioning
Apparent competitors
may actually have
slightly different niches
Species may use
resources in a different
way or time
Minimizes competition
and allows coexistence
Predation
Predators are animals that feed on other
living organisms
Predators are free-living; they do not
take up residence on their prey
Coevolution
Natural selection promotes traits that
help prey escape predation
It also promotes traits that make
predators more successful at capturing
prey
Variation in Cycles
An association in predator and prey
abundance does not always indicate a
cause and effect relationship
Variations in food supply and additional
predators may also influence changes
in prey abundance
Species Interactions
• Canadian lynx
(dashed lines)
versus
snowshoe hares
(solid line)
Prey Defenses
Camouflage
Warning coloration
Mimicry
Predator Responses
Any adaptation that protects prey may
select for predators that can overcome
that adaptation
Prey adaptations include stealth,
camouflage, and ways to avoid
chemical repellents
Parasitism
Parasites drain nutrients from their
hosts and live on or in their bodies
Natural selection favors parasites
that do not kill their host too quickly
Types of Parasites
Microparasites
Macroparasites
Social parasites
Parasitoids
Biological Controls
Parasites and parasitoids are
commercially raised and release in
target areas as biological controls
An alternative to pesticides
Must be carefully managed to not upset
natural balances
Study: Cowbirds
Brown-headed cowbirds evolved in the Great
Plains region of the U.S., living as
commensalists with bison
As they migrated along with the bison, the
cowbirds learned to lay eggs in the nests of
other birds
Cowbird hatchlings push the “host” bird eggs
out of the nest
Today, cowbirds parasitize some 15 species
of birds
End
Succession
Change in the composition of different
of species over time
Pioneer Species
Species that colonize barren habitats - first come,
first served!
Lichens, small plants with brief life cycles
Improve conditions for other species who then
replace them
lose out to better adapted species
who themselves lost out to even better adapted species.
Types of Succession
Primary succession new environments
Secondary succession communities were destroyed or
displaced
Following hurricanes
After the tsunami
Climax Community
Eventually we arrive at the climax community
Stable array of species that persists relatively
unchanged over time
Succession does not always move
predictably toward a specific climax
community; other stable communities may
persist
Multiple outcomes.
Cyclic Changes
DYNAMIC and STABLE
Cyclic, nondirectional changes also
shape community structure
Tree
falls cause local patchiness in tropical
forests
Fires
periodically destroy underbrush in
sequoia forests
Community Instability
REMEMBER
Disturbances can cause a
community to change in ways that
persist even if the change is
reversed
Keystone Species
A species that can dictate community
structure
Removal of a keystone species can
cause drastic changes in a community;
can increase or decrease diversity…
Lubchenco Experiment
QuickTime™ and a
TIFF (Uncompressed) decompressor
are needed to see this picture.
Periwinkles promote or limit diversity in different habitats
Tidepools
Rocks exposed at high tide
Species Introductions
Introduction of a nonindigenous species can
decimate a community
Just recently Hawaii has introduces Sea Stars to
control sea weed
WHY? There are;
No natural enemies or controls
Can outcompete native species
Exotic Species
Species that has left its home natural
range and become established elsewhere
Becomes part of its new community
Can have beneficial, neutral, or harmful
effects on a community…
Kudzu in Georgia
QuickTime™ and a
TIFF (Uncompressed) decompressor
are needed to see this picture.
QuickTime™ and a
TIFF (Uncompressed) decompressor
are needed to see this picture.
Kudzu in Georgia
Quic kTime™ and a
TIFF (Unc ompres sed) decompress or
are needed to see this picture.
Imported for erosion control
No natural herbivores, pathogens, or
competitors
Grows over landscapes and cannot be
dug up or burned out
May
turn out to have some commercial use
for humans
Caulerpa taxifolia
QuickTime™ and a
TIFF (Uncompressed) decompressor
are needed to see this picture.
QuickTime™ and a
TIFF (Uncompressed) decompressor
are needed to see this picture.
Caulerpa taxifolia
Researchers developed a hybrid, sterile
strain of this green alga - for fish tanks.
Somehow it was released into the wild,
where it reproduces asexually - down
the drain!!!
Thrives in varying conditions and emits
a toxin that poisons invertebrates and
fishes
Illegal to import the “aquarium” strain
into the U.S.
Rabbits in Australia
QuickTi me™ and a
TIFF ( Uncompressed) decompressor
are needed to see thi s pi ctur e.
Rabbits were introduced for food and
hunting
Without predators, their numbers
soared
Attempts at control using fences or
viruses have thus far been unsuccessful
QuickTime™ and a
TIFF (Uncompressed) decompressor
are needed to see this picture.
Biodiversity
The sum of all species occupying a
specified area during a specified
interval, past and present
Diversity by Latitude
Diversity of most groups is greatest
in tropics; declines toward poles
a)
ant species
b) breeding birds
Why Are Tropical Species Rich?
Resources are plentiful and reliable
Species diversity is self-reinforcing
Rates of speciation are highest in the
tropics
Surtsey: Volcanic Island
Primary succession
Distance Effect
The farther an island is from a
mainland, the fewer species. WHY?
Closer islands receive more immigrants
Species that reach islands far from
mainland are adapted for long-distance
dispersal and can move on
Distance Effect
Endangered Species
A species that is extremely vulnerable to
extinction
Habitat loss is putting major pressure on
more than 90 percent of the endemic species
that now face extinction
Conservation Biology
Is a systematic survey of the full range
of biological diversity
Entails efforts to decipher biodiversity’s
evolutionary and ecological origins
Attempt to identify methods of
maintaining and using biodiversity in
ways that can benefit the human
population
Only
real way to save the local ecosystem
Sampling of Current Species Diversity
Areas at Risk
Preserving Biodiversity
Requires identifying and protecting
regions that support the highest levels
of biodiversity
It is possible to protect a habitat and still
withdraw resources in a sustainable
fashion - but based on human nature it
is a risky practice!