Transcript Temperature
An Introduction to Ecology
&
the Biosphere
CAMPBELL & REECE
CHAPTER 52
Ecology
from Greek, oikos = home
scientific study of interactions between organisms &
environment
Scope of Ecological Research
Organismal Ecology: concerned with individual’s structure,
physiology, behavior & its challenges posed by its environment
Population Ecology:analyzes factors that affect population
size; how & why it changes over time
Community Ecology: interactions between species: how
predation, competition affect community structure
Ecosystem Ecology: nrg flow & biochemical cycling between
organisms & their environment; abiotic factors included
Landscape Ecology: factors controlling exchanges of nrg,
materials & organisms across multiple rcosystems
Global Ecology: how regional exchange of nrg & materials
influences functioning & distribution of organisms across the
biosphere
Climate
long-term, prevailing weather conditions in given
area
*most significant influence on the distribution of
organisms on land & in oceans
4 Components of Climate
Temperature
2. Precipitation
3. Sunlight
4. Wind
1.
Global Climate Patterns
determined mostly by
input of solar nrg
1.
establishes temp variations
cycles of air & water movement
evaporation of water dramatic latitudinal variations in
climate
2. Earth’s movement in space
Latitudinal Variation in Sunlight Intensity
Earth’s curved shape
causes latitudinal variation
in intensity of sunlight.
because sunlight hits
Tropics (23.5° N and 23.5°
S latitude) most directly,
more heat & light /unit
surface area are delivered
there
@ higher latitudes sunlight
strikes Earth @ oblique
angle so light nrg more
diffuse on Earth’s surface
Global Air Circulation & Precipitation
Patterns
intense solar radiation @ equator initiates global
pattern of air circulation & precipitation
hi temps evaporate water warm, wet air rises
flow toward the poles
air cools precipitation dry air masses descend
@ ~ 30° latitude (N & S)
@~60° latitudes air rises cool precipitation
to poles
Global Air Currents
Wind Patterns
air flowing close to surface creates predictable global
wind patterns
as Earth rotates land near equator moves faster than
that @ poles, deflecting the winds from staying on
vertical path
cooling trade winds blow east west in the tropics
prevailing westerlies blow from west east in
temperate zones
Global Wind Patterns
Climate
Macroclimate: patterns on the global, regional, &
landscape level
Microclimate: very fine localized patterns
Climate patterns can be modified by:
seasonal variations in climate
large bodies of water
mountain ranges
Seasonality
Earth’s tilted axis of rotation & revolution around
Sun every year cause strong seasonal cycles in mid to
hi latitudes
Bodies of Water
because of hi specific heat of water, oceans & large
lakes tend to moderate the climate of nearby land
hot day: land warmer than water air over land
warms & rises draws cooler air from over water to
land
@ night: land cools faster than water air over now
warmer water rises draws cooler air over land
back over water
Lake-Effect Snow
Global Circulation of Surface Water
Mountains
Microclimate
every environment on Earth is characterizes by
small-scale differences in abiotic factors
chemical & physical attributes:
temperature, amt of shade, light, water &
nutrients, fallen tree used as shelter
Global Climate Change
increasing greenhouse gas concentrations in the air
are warming Earth & altering the distributions of
many species
some will thrive
others will not be able to shift their ranges
quickly enough to reach suitable habitat
Biomes
major life zones characterized by vegetation type (in
terrestrial biomes) or by the physical environment
(in aquatic biomes)
Climograph
plot of annual mean
temperature &
precipitation in a
particular region
Climograph for Some Major Biomes
Climographs
show that temp & precipitation are correlated with
biomes
because other factors also play a role in biome
location: biomes can overlap
General Features of Terrestrial Biomes
most named for major physical or climatic features &
for their predominant vegetation
each biome also characterized by:
microorganisms
fungi
animals
all adapted to that particular environment
Ecotone
area of integration: where biomes overlap
Terrestrial Biomes
layering w/in biome due to shapes & sizes of plants
flora dependent on annual precipitation & temps
Biome Species Composition
varies w/in each biome
ex: eastern part of one large lake may have
different water bird than western portion
Disturbance
event that changes a community: removes organisms
from it & alters the resource availability
ex: forest fire
Tropical Forest
Distribution: equatorial & subequatorial
Precipitation:
Tropical
Rainforest: constant, 200 -400 cm/yr
Tropical Dry Forest: seasonal, 150 – 200 cm/yr
Temperature:
high all yr, average 25 – 29°C , little seasonal
variation
Tropical Forest
Tropical Forest: Plants
vertically layered
intense competition for light
Tropical Forest Plants
Tropical Dry Forest
Tropical Rainforest
see all layers, some
with 2 layers of
subcanopy trees
broadleaf evergreen
trees dominate
epiphytes (air
plants) & orchids
typically cover trees
see fewer layers
drop leaves during
dry season
commonly have
thorny shrubs &
succulent plants
Tropical Forest: Animals
millions of species
5 – 30 million undiscovered species of insects,
spiders, other arthropods
highest animal diversity than anywhere else on Earth
all adapted to vertically layered environment
Tropical Forest: Human Impact
thriving communities of man have lived in tropical
forests for hundreds of years
overpopulation leading to agriculture & development
are destroying many tropical forests
DESERT
Distribution:
occur in bands near 30° N & S latitude or in
interior of continents
Precipitation:
low & variable; <30 cm/yr
Temperature :
variable seasonally & daily
hot desert: max T may > 50°C
dry desert: low T may < -30°C
World Distribution of Deserts
Deserts
Desert Plants
see low, widely scattered vegetation
see more bare ground than other terrestrial
biomes
succulents
cacti
euphorbs
deeply rooted shrubs & herbs
grow during brief rainy periods
Desert Plants
Adaptations:
heat & desiccation tolerance
water storage
reduced leaf surface area
CAM photosynthesis
physical defenses:
spines
chemical defenses:
toxins in leaves of shrubs
Desert Animals
Common animals:
Snakes
Lizards
Scorpions
Ants
Beetles
Birds: migratory & resident
seed-eating Rodents
Desert Animal Adaptations
many species are nocturnal
water conserved in variety of ways:
only water some get is by metabolizing
carbohydrates water + carbon dioxide
Desert: Human Impact
use of long distance transport of water & deep
groundwater wells have allowed large populations of
man to make the desert their home
end result decreased diversity of some deserts
SAVANNA
Distribution:
equatorial & subequatorial
Precipitation:
seasonal rainfall 30 – 50 cm/yr
dry season can last 8 – 9 months
Temperature :
warm year-round: 24 – 29 °C
more seasonal variation than tropical forests
Savanna Distribution
Savanna
Savanna Plants
scattered, variable density of trees
most plants have small leaves (adaptation to dry
conditions)
Fires common in dry season: most dominant plant
species are fire-adapted & drought-tolerant
grasses & forbes (clover, wildflowers) tolerant of
large grazing herbivores
Savanna Animals
dominant herbivores are insects
especially termites
large herbivores migrate toward thicker vegetation &
watering holes during dry season
Savanna: Human Impact
earliest humans lived in the savanna
agriculture & hunting (poaching) have reduced #s of
large mammals
Chaparral
also called
mattoral (Spain & Chile)
garigue & maquis (southern France)
fynbos (South Africa)
Chaparral
Distribution:
midlatitude coastal regions
Precipitation:
highly seasonal (rainy winters, dry summers)
averages 30 – 50 cm/yr
Temperature :
fall, winter, spring are cool (10 – 12°C)
summer can get > 40°C
Chaparral Distribution
Chaparral
Chaparral Plants
dominated by shrubs, small trees, variety of grasses
& herbs
plant diversity high though some species found only
in very limited areas
adaptations to:
drought: tough evergreen leaves
fire:
herb seeds only germinate after hot fire
roots are fire resistant (plants re-sprout quickly)
Chaparral Animals
natives include:
browsers (deer,
goats)
high diversity of
small mammals
many amphibians,
birds, reptiles,
insects
Chaparral: Human Impact
due to increased agricultural use of land chaparral
areas have been heavily settled & reduced
man contributes to fires
Temperate Grassland
also called:
veldts (South Africa)
puszta (Hungary)
pampas (Argentina & Uruguay)
steppes (Russia)
plains & prairies (North America)
Temperate Grasslands
Temperate Grassland
Precipitation:
highly seasonal: dry winters/wet summers
averages vary between 30 – 100 cm/yr
periodic drought is common
Temperature :
winters cold (< -10°C)
summers moderately hot ( 30°C)
Temperate Grasslands
Temperate Grasslands: Plants
dominant plants are grasses & forbs
some grasses 2 m high
many adapted to survive periodic drought & fires
grazing by herbivores helps prevent establishment of
woody plants
Temperate Grasslands: Animals
native mammals
large: bison, wild horses
small burrowers: prairie dogs
Temperate Grasslands: Human Impact
most grasslands of North America & Eurasia
converted to farmland
in other grasslands grazers have turned the
grasslands deserts
desertification: Patagonia, Argentina
Northern Coniferous Forest
aka: taiga
Distribution:
broad band across northern North America &
Eurasia to edge of arctic tundra
Precipitation:
30 – 70 cm/yr
periodic droughts are common
Temperature :
winters cold (-50°C in Siberia)
summers usually >20°C
Northern Coniferous Forest
Northern Coniferous Forest: Plants
dominated by cone-bearing trees
pine, spruce, fir, hemlock
some require fire to regenerate
shape of conifers prevents too much snow
accumulating…so branches don’t break
needle-or scale-like leaves reduce water loss
lower diversity of shrubs & herbs than in temperate
broadleaf biomes
Northern Coniferous Forest: Plants
Northern Coniferous Forest: Animals
Birds: residents & summer migrants
insects occasionally kill large tracts of trees
Mammals:
Moose
Brown Bear
Siberian Tiger
Northern Coniferous Forest: Human
Impact
logging increasing at alarming rate
not many old stands remain
Temperate Broadleaf Forest
Distribution
mainly in midlatitudes of northern hemisphere
smaller areas in Chile, South Africa, Australia,
New Zealand
Temperate Broadleaf Forest
Precipitation:
70 to > 200 cm/yr (includes snow)
all seasons have precipitation
Temperature :
winter averages ~ 0°C
summers hot & humid/ up to 35°C
Temperate Broadleaf Forest
Temperate Broadleaf Forest: Plants
mature forest has distinct vertical layers including a
closed canopy
dominant plants in North America are deciduous
trees
adaptation: drop leaves as weather gets colder:
uptake of water by roots not feasible when soil
frozen
dominant plant in Australia: Eucalyptus
Temperate Broadleaf Forest: Animals
mammals, birds, insects make use of vertical layers
many mammals hibernate in winter
many birds (and some butterflies) migrate south
Temperate Broadleaf Forest:
Human Impact
virtually all original deciduous forests in North
America have been destroyed by urban development
or logging…but have great capacity for recovery:
some areas are returning over much of their original
range
Tundra
Distribution:
covers arctic: 20% Earth’s land surface
tops of high mountains
Precipitation:
20 – 60 cm/yr in arctic tundra
>100 cm/yr alpine tundra
Temperature:
winter averages < -30°C
summer averages < 10°C
Tundra
Tundra
Tundra: Plants
mostly herbaceous:
mosses, grasses,
forbs + dwarf shrubs
& trees, lichens
permafrost (frozen
ground year round)
prohibits growth of
plant roots
Tundra: Animals
Birds: migratory, arriving for nesting in summer
Mammals:
Residents: musk ox
Migrators: caribou, reindeer
Predators: bears, wolves, foxes
Tundra: Human Impact
sparsely populated but has been greatly impacted by
mineral & oil extraction
Aquatic Biomes
charaterized primarily by their physical environment
rather than be climate
often layered with regard to
light penetration
temperature
community structure
Zonation in Aquatic Biomes
light absorbed by water itself + photosynthetic
organisms so…light intensity decreases rapidly with
depth
Photic Zone: sufficient light for photosynthesis
Aphotic Zone: little light penetrates
Pelagic Zone = photic zone + aphotic zone
Zonation in Aquatic Biomes
Abyssal Zone:
2,000 – 6,000 m deep
Benthic Zone:
the bottom of all aquatic biomes, shallow or deep
Benthos:
communities of organisms that live in sand &
sediments of the benthic zone
More Definitions
Detritus:
dead organic material that “rains” down from
photic zone; food source for benthos
Thermocline:
narrow layer of water where there is an abrupt
temperature change
separates the more uniformly warm upper layer
from the uniformly cold deeper water
many temperate lakes undergo a semiannual
mixing of their water
Lakes
lake environment generally classified on basis of 3
physical criteria:
1. light penetration
photic / aphotic
2. distance from shore / depth of water
littoral / limnetic
3. open water / bottom
pelagic / benthic
Marine Zonation
classified by 3 criteria:
light penetration
1.
photic / aphotic
2. distance from shore / depth of water
intertidal / neritic / oceanic
3. open water / bottom
pelagic / benthic / abyssal
Lakes
standing bodies of water range from ponds a few
square meters in area to lakes covering thousands of
square kilometers
Lake: Chemical Environment
lakes differ greatly in their salinity, O2
concentration, & nutrient content
Oligotrophic Lakes:
nutrient poor
O2 rich
low in amt of decomposable matter
Eutrophic Lakes:
nutrient rich
O2 poor in deepest zones in summer
high amt decomposable matter
Lakes: Geologic Features
oligotrophic lakes can become more eutrophic over
time as runoff adds sediments & nutrients
oligotrophic lakes tend to have less surface area
relative to their depth
Lakes: Oligotrophic
Lakes: Eutrophic
Lakes: Photosynthetic Organisms
Littoral Zone:
shallow, well-lit waters close to shore
rooted & floating aquatic plants
Limnetic Zone:
waters too deep to support rooted plants
phytoplankton, including cyanobacteria
Phytoplankton
Lakes: Heterotrophs
Limnetic Zone:
small, drifting heterotrophs or zooplankton
(graze on phytoplankton)
Benthic Zone:
assorted invertebrates (species depends on O2
content)
Fishes live in all zones that have sufficient O2
Zooplankton
Lakes: Human Impact
Runoff from fertilized land & dumping wastes
water nutrient enrichment algal blooms O2
depletion fish kills
Wetlands
habitat that is inundated by water (at least part of
the year) & supports plants adapted to watersaturated soil
due to high organic production by plants &
decomposition by microbes: water & soil of wetlands
periodically low in dissolved O2
*high filter capacity: both nutrients & pollutants
Wetlands: Geologic Features
Basin Wetlands:
develop in shallow basins
range: upland depressions filled in lakes
Riverine Wetlands:
along shallow & periodically flooded banks of
streams
Fringe Wetlands:
along coasts of large lakes & seas
water flows back/forth due to changing water
levels or tides
fresh water & marine biomes
Basin Wetlands
Riverine Wetlands
Fringe Wetlands
Wetlands: Autotrophs
among most productive biomes in world
water-saturated soils great for plants
Lily pads
Cattails
Sedges
Tamaracks
Black spruce
Wetlands: Heterotrophs
diverse community of invertebrates, birds, reptiles,
amphibians, and mammals
Herbevores:
crustaceans
aquatic insect larvae
muskrats
Carnivores:
dragonflies
frogs
alligators
herons
Wetlands: Human Impact
draining & filling have destroyed up to 90% of
wetlands
Streams: Physical Environment
most prominent characteristic: their current
stratified into vertical zones
Streams: Physical Environment
Headwaters:
generally
cold, clear
turbulent, & swift
Downstream:
generally
warmer
more turbid
Streams: Chemical Environment
salt & nutrient concentrations increase as get further
from headwaters
Headwaters: generally rich in O2
Downstream: + O2 unless has organic enrichment
Streams: Geologic Features
headwaters:
downstream:
often narrow with rocky
wide stretches
bottom
alternate between
shallow sections &
deeper pools
meandering
silty bottoms
Streams: Photosynthetic Organisms
rivers that flow thru grasslands or deserts have
phytoplankton or rooted aquatic plants
Streams: Heterotrophs
great diversity of fishes & invertebrates inhabit
unpolluted streams
distributed
in vertical zones
organic matter from terrestrial vegetation is primary
source of food for aquatic consumers
Streams: Human Impact
pollutants from municipal, agricultural, & industrial
sources kill aquatic organisms
damming & flood control impair natural functioning
of stream ecosystems & threaten migratory species
(salmon)
Estuary
a transitional area between river & sea
when high tide: salt water flows up estuary channel
higher density sea water stays below lesser density
freshwater
Estuary: Chemical Environment
salinity varies from that of freshwater sea water &
with rise & fall of tides
nutrients from rivers make estuaries some of most
productive biomes
Estuary: Geologic Features
complex network of tidal channels, islands, natural
levees, & mudflats
Estuary: Photosynthetic Organisms
saltmarsh grasses & algae (including phytoplankton)
are major producers
Estuary: Heterotrophs
abundant #’s of worms, oysters, crabs, & many fish
many invertebrates & fishes use estuaries as
breeding grounds
crucial feeding grounds for birds & some marine
mammals
Estuary: Human Impact
Filling, dredging, & pollution have disrupted
estuaries worldwide
Intertidal Zones
are periodically submerged & exposed by the tides,
2x daily on most marine shores
upper zones exposed to air for longer periods
greater variation in temp & salinity
changes in physical conditions from upper to lower
zones limits the distribution of many organisms to
particular strata
Intertidal Zones: Chemical Environment
O2 & nutrient levels generally high & renewed with
each turn of the tides
Intertidal Zone: Photosynthetic
Organisms
high diversity & biomass of attached marine algae
inhabit rocky intertidal zones
much lower diversity & biomass in sandy intertidal
zones with vigorous wave action
sandy intertidal zones in protected bays or lagoons
have rich beds of grass & algae
Intertidal Zone: Heterotrophs
animals here have multiple structural adaptations
rocky
areas: ways to attach to hard surfaces
sandy areas: many bury themselves
feed on what tides bring them
Intertidal Zones: Human Impact
oil spill have disrupted ecosystem of many intertidal
zones
construction of rock walls, barriers to reduce damage
from erosion, storm surges also disrupts these zones
Ocean Pelagic Zone
open blue waters
mixed constantly by wind & ocean currents
photic zone extends deeper here (water is clearer)
Oceanic Pelagic Zone: Chemical
Environment
O2 levels generally high
nutrient levels generally lower than in coastal waters
tropical oceans: thermally stratified all year
temperate & hi-latitude oceans have spring & fall
turnover so generally nutrients renewed in photic
zone
Oceanic Pelagic Zone: Geologic Features
covers ~70% Earth’s surface
average depth = 4,000 m
deepest point: 10,000 m
Pelagic Zone: Photosynthetic Organisms
phytoplankton (including photosynthetic bacteria)
dominate
due to vast area this zone covers: ~50% of all
photosynthesis on Earth by them
Pelagic Zone: Heterotrophs
zooplankton most abundant group in this zone
graze on phytoplankton
includes:
protists
worms
copepods
shrimp-like
krill
jellies
small
larvae of invertebrates
Pelagic Zone: Heterotrophs
also include free-swimming animals:
large
squid
fishes
sea turtles
marine mammals
Pelagic Zone: Human Impact
overfishing has depleted fish stocks in all oceans
all also polluted
Coral Reefs
formed largely from the calcium carbonate skeletons
of corals
in photic zone of relatively stable tropical marine
environments with high water clarity
sensitive to temps < 18 – 20° & > 30°C
found in deep seas 200 -1,500 m deep
as
much diversity as shallow reef
Deep Sea Coral Reef
Shallow Coral Reef
Coral Reef: Chemical Environment
require high O2 levels
Coral Reef Geologic Features
Corals require a solid substrate for attachment
typically: begins as fringing reef on young, high
island forming an off-shore barrier reef as island
ages coral atoll
Barrier Reef
Coral Atoll
Coral Reefs: Photosynthetic Organisms
unicellular algae live w/in tissues of corals in
mutualistic relationship: provides corals with
organic molecules
diverse multicellular red & green algae growing on
reef also photosynthesize
Coral Reef: Heterotrophs
dominant
heterotroph: corals
are a diverse group of
cnidarians
also high diversity of
fishes & invertebrates
overall nearly as
diverse as tropical
rainforest
Coral Reef: Human Impact
populations of corals & fishes on decline due to
humans collecting corals & overfishing
Global warming & pollution coral death
Marine Benthic Zone
consists of the seafloor under surface waters of the
coastal (neritic) zone * the offshore (pelagic) zone
Benthic Zone
near-coastal areas only part to receive sunlight
water temp declines with depth while pressure
increases
organisms
in very deep abyssal zone adapted to cold
(~3°C) & high water pressure
Benthic Zone: Chemical Environment
O2 levels usually high enough to support divers
animal life
Benthic Zone: Geologic Features
most covered by soft sediments
also: rocky surfaces, submarine mts, new oceanic
crust
Benthic Zone: Autotrophs
shallow areas: seaweeds & filamentous algae
deep-sea hydrothermal vents:
on
mid-ocean ridges
chemo-autotrophic prokaryotes obtain nrg by oxidizing
H2 S formed by a reaction between hot water &
dissolved sulfate (SO4 )
Benthic Zone: Heterotrophs
numerous invertebrates & fishes
beyond photic zone most animals rely on organic
material raining down from above
many around hydrothermal vents
Giant tube worms: eat chemo-autotrophic
prokaryotes that live as symbionts w/in their bodies
Benthic Zone: Human Impact
overfishing has decimated some benthic fish
populations (cod)
dumping of organic wastes has created oxygendeprived benthic areas
Distribution of Species
are a consequence of both ecological & evolutionary
interactions thru time
Ecological Time
differential
survival & reproduction of individuals that
lead to evolution
Evolutionary Time
thru
natural selection, organisms adapt to their
environments over time frame of many generations
Global & Regional Patterns
ecologists ask not only where a species lives but also
why it lives there
to
answer these ?s focus on both biotic & abiotic factors
that influence distribution & abundance of organisms
Flowchart of Factors Limiting Geographic
Distribution
Dispersal
movement of individuals or gametes from their areas
of origin or from centers of high population density
dispersal of organisms is critical to understanding
the role of geographic isolation in evolution
Natural Range Expansion
long distance dispersal can lead to adaptive
radiation: the rapid evolution of ancestral species
into new species that fill many ecological niches
Species Transplants
by observing the results of intentional or accidental
transplants of species to areas where it was
previously absent, ecologists may determine if
dispersal is a key factor limiting distribution of a
species
species introduced to new geographic locations often
disrupt the communities & ecosystems to which they
have been introduced & usually spread beyond area
of introduction
Behavior & Habitat Selection
habitat selection one of least understood processes
Insect: some females will only lay eggs near plant
that species most prefers thus limiting habitat
Biotic Factors
If behavior does not limit distribution of species then
do other species influence it?
Often (-) interactions with predators or herbivores
restricts ability of a species to survive & reproduce
Biotic Factors
besides presence or absence of predators or
herbivores presence or absence of pollinators, food
resources, parasites, pathogens, & competing
organisms can act as biotic limitations on
distribution of a species
Abiotic Factors
temperature, climate, water, oxygen, salinity,
sunlight, soil can all limit a species distribution
most areas have fluctuations in nearly all these
abiotic factors
some organisms can avoid some of the more extreme
annual fluctuations
dormancy
storage
of food or water supplies
Temperature
important abiotic factor in distribution of organisms
because of its effect on biological processes
cells would rupture if water in them freezes when
0°C
proteins of most organisms would denature
if temp > 45°C
extraordinary adaptations allow some species to
survive in temp ranges other organisms cannot
survive in
Colors of Pool due to Thermophiles
Water & Oxygen
terrestrial organisms face nearly a constant threat of
dehydration: their distribution reflects their ability
to obtain & conserve water
water affects oxygen availability in aquatic
environments & in flooded soils
surface waters of streams tend to be well oxygenated
due to rapid exchange with atmosphere
Salinity
affects water balance of organisms thru osmosis
most organisms can excrete excess salts by
specialized glands or in feces & urine
salt flats or other high-salinity habitats have few
species of plants or animals
Salmon go from freshwater salt-water & back have
both behavioral & physiological mechanisms to
osmoregulate
they
adjust amt water they drink to balance their salt
content
gills switch from taking up salt in freshwater to
excreting salt in sea-water
Sunlight
in aquatic environments, every meter of water depth
absorbs 45% of red light & ~2% of blue light passing
thru it
Sunlight
too much increases temps as in deserts which
stresses plants & animals
@ high elevations sunlight more likely to damage
DNA & proteins because atmosphere is thinner so
get more UV radiation
this damage + other abiotic factors reason why there
is a tree line on mountain slopes
Rocks & Soil
pH, mineral composition, physical structure of rocks
& soil limit distribution of plants & therefore animals
that feed on them
pH can act directly thru extreme acidic or basic
conditions or indirectly by affecting the solubility of
nutrients & toxins
composition of riverbeds can influence water
chemistry influences organisms that can live there