Transcript SHOW-6-2005

Cell Walls
Cell walls are:
– Structural
– Provide defense against invading
pathogens
– Provide signaling pathways for cells
Cell Wall Chemistry
• Cell walls contain cellulose, pectin
and lignin
• Cellulose is the major component
– Most abundant polymer in the world
~40-60% of dry weight of plants
– Form strong strands of parallel chains
called Cellulose Microfibrils
– Cellulose strands are cemented
together by pectins
TEM photograph
Pectin and Lignin
• Pectins are polymers of a-galacturonic
acid
– Flexible, long molecules that link
Microfibrils together…like ‘glue’
• Lignin is very hard, tough, rigid
– Chemistry is not fully elucidated
– Polymer of complex, amino-acid derived
monomers
– Creates strong cell walls
And don’t forget the 5 colors !
Plant Tissues
• Tissues are groups of cells that form
a structural and functional unit
• Vascular plants have three tissue
systems:
– Dermal tissue
Outer covering for the plant body
– Ground tissue
Photosynthesis, storage and support
– Vascular tissue
Water and nutrient conduction system
Functions for Plant
Epidermis
1) Prevent water loss in
shoot (waxy surface
and bark)
2) Controlling exchange
of gases in shoot
(stomates)
3) Absorption by roots
(root hairs)
4) Produce hairs, called
trichomes, that shade
the leaf or protect from
herbivores (physically
or chemically)
A
Gymnosperm
B
Dicot
C
Monocot
Yam
D
Monocot
E
Monocot
Do Animals have epidermis?
What is animal epidermis called?
What does this have to do with Safe Sex?
Cheek cells
Jicama -- Pachyrhizus erosus (L.)
Jicama -- Pachyrhizus erosus (L.)
Eaten in celebration of
Day of the Dead
November 1st
Mexico
Plant Tissues
• Tissues are groups of cells that form
a structural and functional unit
• Vascular plants have three tissue
systems:
– Dermal tissue
Outer covering for the plant body
– Ground tissue
Photosynthesis, storage and support
– Vascular tissue
Water and nutrient conduction system
Vascular Tissues - Xylem
• Xylem conducts water and nutrients from
roots to the rest of the plant
• Two types
– Tracheids
• Dead at maturity
• Main water-conducting cells of fern and gymnosperms
• Relatively few plasmodesmata in end of cell in pit fields,
thin areas w/only primary cell wall
– Vessel elements
• Dead at maturity – stacked on top of each other
• Broader than tracheids, large holes in ends of cell
• Much more efficient at water transport
End wall with
perforations
Xylem carries
water &
soil nutrients
Pits
Cell wall
Lumen
Tracheids
Vessels
(a)
(b)
Xylem Vessel
Xylem
Tracheid
Vascular Tissues - Phloem
• Transports food materials
• Provides structural support
• Two types:
- Sieve tube cells
• Alive at maturity
• Extensive holes via sieve plate at cell ends, with shared
cytoplasm between cells
• Reduced cellular organelle volume – some lose nucleus
– Companion cells
• Nucleated, support cells for sieve tubes
• Connected by many plasmodesmata to sieve tube cell
• Involved in sugar transport to sieve tube cell
Sieve plate
Sieve tube
member
Phloem
parenchyma
cells
Lateral sieve
area
Plasmodesma
Companion cell
(c)
(d)
Fig. 31.05cd
Companion Cell
Sieve Tube
Cell
Xylem and Phloem Cells
Fiber Cells
Xylem & Phloem: (in Squash)
Fibers
Phloem
Xylem
Xylem
Vascular Cambium
Epidermis
Phloem
Vascular
Cambium
Xylem
Notice that the
vascular cambium
circles all the way
around
Fibers
Dicot
stem
Hemp
Jute for burlap
Examples of plant fibers
from stem vascular bundles
Sisal = fiber from
monocot stem
Epidermis
Vascular
Bundles
(scattered)
Monocot Stem
Monocot Stem
Air Space
Xylem (big cells &
smaller ones in
circle
Phloem (all cells
within box
Secondary Growth
Secondary Growth……Why?
* Plant is getting bigger so needs more materials
* Old vascular tissue gets “clogged up”
1, 2, 3 years
Of age
Pith
Rays in wood
Tree Borer
For assessing tree rings
Bristlecone Pine
How old are these trees?
Oldest one is
4,800 years old
The oldest living trees on earth.
The Curly Redwood Lodge is one of northern California’s most
unique lodges. It was built from one curly redwood tree that produced
57,000 board feet of lumber. The tree (cut down in 1952) was 18 feet
wide at the trunk. Curly redwood is unique because of the curly grain
of the wood, unlike typical straight grained redwood.
http://www.curlyredwoodlodge.com/rooms.htm
1952
Jewelry meant for Queen Nerfititi (Egypt) was
found on a sunken wooden ship that was built
in 1316 B.C.
King Midas (Turkey) was buried in a funeral
mound built with logs that were cut in 718 B.C.
Lost Colony of Roanoke, Virginia
Can also study when major forest fires occurred in the past.
A researcher in Tennessee noticed that
hurricanes cause heavy-oxygen to
accumulate in “late wood” (because
hurricanes usually occur in late fall).
She has been accurate back to 100
years ago….wants to go back 500 years.
• Two small, old-mining towns that are currently
military posts (Arizona and Nevada)
• Unusually high levels of childhood leukemia cases
• High levels of tungsten in the urine of townspeople
• Is there a cause:effect relationship between tungsten (a heavy
metal, like lead or mercury) exposure and leukemia? …still being
investigated
• But tree-ring specialists have discovered that the levels
of tungsten in the environment have increased dramatically
in the last 20 years….each annual ring captured whatever
chemicals that might have occurred in the environment that
year!!!
September, 2002
Sapwood
Heartwood
Sycamores
Cork
Fibers
Phloem
Xylem
Girdling can kill plants
Carpenters talk about two kinds of wood:
Softwood and Hardwood
Softwood = light, easy to put nails/screws into…so are best
for construction. Gymnosperms generally make
softwoods. Since Pines grow so fast they are the
most popular. Paper is made from pine mostly.
Hardwood = from Dicot trees mostly; more of a mixture of
xylem cells (including fibers); that makes them
more dense. Used for furniture and art/crafts.
Gymnosperms like pines,
junipers, spruce, fir, and
redwood.
Most of the xylem cells are
one kind...called Tracheids.
Also, little or no Fibers.
Dicots like oak, maple,
ash, hickory, walnut.
Contain both tracheids and
vessels and fibers.
The fibers make the wood
harder to cut and nail into,
But will also be sturdier for
Making furniture, flooring,
art, etc.
Not all Hardwoods are
actually “harder” than
Softwoods, and not all
Softwoods have wood that
is “softer” than Hardwoods.
Hickory
(Hardwood)
•Wright Brothers plane
•Pioneer wagon wheels
•Baseball bats
• Skis
•Axes, hammer handles, etc.
• Golf clubs
• wooden floors
Mostly tracheids
& fibers with
A very large
vessels
interspersed.
Very slow
growing.
Some nonvascular cells
are gelatinous
of Wheat
Starchy food supply
(Endosperm)
Dietary Fiber
Scutellum = Cotyledon
Embryo
It is believed that the early hominids were
eventually forced to live in a biome that
was different than the Tropical Rainforest.
This might have been due to: 1) globalclimate change, or 2) rising competition for
resources due to population pressures, or
3) a combination of these two factors.
Which biome do you think they would
have ended up in (according to this
scenario)?
Primates typically live in Tropical
Rainforests (using hands & feet to climb
trees)….
…they eat leaves, fruits, seeds & insects.
< Living in the Savanna??
Primates typically live in Tropical
Rainforests (used hands & feet to climb
trees)….
…eats leaves, fruits, seeds & insects.
Chimpanzee (5 million years ago)
…eats leaves, fruit, seeds, resin, bugs
Australopithecus (3 million years ago)
...eating more roots, grains & meat
Roots, tubers, bulbs, rhizomes
Savanna
Tropical Rainforest
More difficult life…more challenges:
• Bipedalism (to move further & to see
predators & prey better & get less sun
& use hands to carry things
• Develop use of tools
• More social organization (language?)
• Larger brains
• Larger teeth to eat roots & grains & meat
< Climate-change…more Savanna-like
Primates typically live in Tropical
Rainforests (used hands & feet to climb
trees); eat leaves, fruits, and insects.
Taro root
Carrots
Cassava
Parsnips