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The Nervous System
Chapter 26
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Outline
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Invertebrate Nervous System
Neurons
Nerve Impulses
Synapse
Drug Addiction
Evolution of Vertebrate Brain
Vertebrate Nervous System
Sensory Receptors
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Evolution of Animal Nervous System
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In most invertebrates, and all vertebrates,
sensory receptors and motor effectors are
linked by the nervous system.
Association neurons are located in the
brain and spinal cord, and together are
know as central nervous system.
Motor neurons and Sensory neurons
together comprise peripheral nervous
system.
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Three Types of Neurons
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Invertebrate Nervous Systems
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Sponges are the only major phylum of
multicellular animals lacking nerves.
Reflexes - Cnidarian neurons are linked
together through nerve net.
No associative activity.
Associative Activities - Free-living flatworms
have two nerve cords running down bodies.
Permits complex control of muscles.
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Invertebrate Nervous Systems
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Evolutionary Path to Vertebrates
More sophisticated sensory mechanisms.
Differentiation into central and peripheral
nervous systems.
Differentiation of sensory and motor
nerves.
Increased complexity of association.
Elaboration of the brain.
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Evolution of Nervous System
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Neurons
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All neurons have same basic structure.
Cell Body is enlarged region containing
nucleus.
Dendrites are short slender input channels
extending from end of cell body.
Axon is single output channel extending
out from other end of cell body.
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Neurons
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Most neurons require nutritional support
provided by companion neuroglial cells.
Schwann cells and oligodendrocytes
envelop axon with sheath of myelin acting
as electrical insulator.
- During development, cells wrap
themselves around each axon several
times to form myelin sheath.
Uninsulated gaps (nodes of Ranvier).
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Neuron Structure and Myelin Sheath Formation
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The Nerve Impulse
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When a neuron is at rest, active transport
channels in plasma membrane transport
sodium ions out of the cell and potassium
ions into the cell.
Plasma membrane is polarized.
- Resting Potential is charge separation
between outside and inside of cell.
- Sodium channels flood into neuron from
outside, momentarily depolarizing inside
of membrane.
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The Nerve Impulse
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Electrical current moves in and out of neuron
through voltage-gated channels.
Sodium channels flood into neuron from
outside, momentarily depolarizing inside of
membrane.
- Moving reversal known as action
potential.
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The Synapse
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Synapse is junction of axon with another cell.
Presynaptic Membrane is located on near
side of synapse.
Postsynaptic Membrane is located on far
side of synapse.
Neurotransmitters are chemical messengers
that carry nerve impulses across synapse.
Control switches of nervous system.
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Synapse Events
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The Synapse
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Excitatory Synapse - Sodium channel
through membrane is opened by chemical
neurotransmitter.
Inhibitory Synapse - Binding with
neurotransmitter opens potassium channel
and inhibits start of an action potential.
Integration - Various excitatory and
inhibitory electrical effects cancel or
reinforce one another.
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Drug Addiction
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When a cell is exposed to chemical signal for
a prolonged period, it tends to lose ability to
respond with its original intensity.
If receptor proteins within synapses are
exposed to high levels of neurotransmitter
molecules for prolonged periods, the nerve
cell often responds by inserting fewer
receptor proteins into the membrane.
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Drug Addiction
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Cocaine
Neuromodulator (prolongs transmission of
signal across synapse) that causes large
amounts of neurotransmitters to remain in
synapses for long periods of time.
- Transmit pleasure messages using the
neurotransmitter dopamine.
Nerve cells may eventually lower
number of receptor proteins on
surface.
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Evolution of the Vertebrate Brain
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Hindbrain was major component of early
brains, and is still found in fishes today.
May be considered extension of spinal
cord devoted primarily to coordinating
motor reflexes.
- Much of coordination carried on within
cerebellum.
Remainder of brain devoted to
reception and processing sensory
information.
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Evolution of the Vertebrate Brain
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Midbrain is composed primarily of optic lobes
which receive and process visual information.
Forebrain is devoted to processing olfactory
information.
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Evolution of the Vertebrate Brain
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Dominant Forebrain
Sensory information increasingly centered in
forebrain starting with amphibians.
- Diencephalon
Thalamus - Relay center between
incoming information and cerebrum.
Hypothalamus - Controls pituitary
secretions.
- Telencephalon - Associative activity
(Cerebrum)
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Evolution of Vertebrate Brain
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How the Brain Works
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Cerebrum is large rounded area of brain
divided by groove into right and left halves.
Functions in language, conscious thought,
memory, personality, vision, and a host of
other activities.
- Much of activity occurs in thin gray outer
layer (cerebral cortex).
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Human Brain
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How The Brain Works
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Thalamus is major site of sensory
processing, and controls balance.
Hypothalamus integrates all internal
activities.
Controls centers in brain stem regulating
body temperature, blood pressure,
respiration, and heartbeat.
Linked to areas of cerebral cortex by limbic
system.
- Deep-seated drives
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How The Brain Works
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Language and Other Higher Functions
Left hemisphere is dominant hemisphere
for language.
- Adept at sequential reasoning.
Non-dominant hemisphere is adept at
spatial reasoning, musical ability, and
important for consolidation of memories of
nonverbal experiences.
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The Spinal Cord
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Cable of neurons extending from the brain
down through the backbone.
Surrounded and protected by vertebrae,
through which spinal nerves pass out to
the body.
- Motor nerves from spine control most
muscles below the head.
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Vertebrate Nervous System
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Voluntary Nervous System
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Voluntary Nervous System relays commands
to skeletal muscles.
Can be controlled by conscious thought.
Reflex - Produces rapid motor response to
stimulus because sensory neuron passes
information directly to a motor neuron.
Most involve single connecting interneuron
between sensory neuron and motor
neuron.
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Autonomic Nervous System
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Autonomic Nervous System stimulates glands
and relays commands to smooth muscles.
Cannot be controlled by conscious thought.
Composed of:
- Sympathetic nervous system - Takes
command in time of stress.
- Parasympathetic nervous system Conserves energy by slowing processes
down.
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Sensory Receptors
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Sensory receptors are specialized sensory
cells that detect changes inside and outside
the body.
Sensory organs - relatively complex
receptors.
Path of sensory information
Stimulation
Transduction
Transmission
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Sensory Receptors
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Receptors can initiate nerve impulses by
opening or closing stimulus-gated channels.
Exteroceptors - Sense stimuli in external
environment.
Interoceptors - Sense stimuli within the
body.
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Sensing the Internal Environment
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Vertebrate body uses variety of sensory
receptors to respond to internal environment.
Temperature Change
Blood Chemistry
Pain
Muscle Contraction
Blood Pressure
Touch
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Sensing Gravity: Balance
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Brain needs point of reference to maintain
balance.
Otolith sensory receptors are located in
hollow chambers within inner ear and
detect gravity.
Brain senses motion by employing a receptor
in which fluid deflects cilia in a direction
opposite that of motion.
Semicircular canals
- Cilia arranges in cupula assembly.
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Inner Ear and Motion Sensing
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Sensing Chemicals: Taste and Smell
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Taste
Taste buds
- Papillae
Chemicals dissolve in saliva and
contact taste cells.
Smell
Chemically sensitive neurons in nose have
cell bodies embedded within epithelium of
the nasal passage.
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Taste and Smell
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Sensing Sounds: Hearing
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Eardrum membrane pushed in and out by
waves of air pressure.
Three small bones located on other side of
eardrum act as lever system to increase
vibration force.
- Inner ear chamber (cochlea) connected
to throat by eustachian tube so that
pressure is equalized.
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Sensing Sounds: Hearing
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Sound receptors within cochlea are hair cells
resting on a membrane running up and down
middle of chamber.
When sounds enter cochlea, sound waves
cause membrane to vibrate, causing hairs
to send nerve impulses to sensory neurons
that travel to the brain.
- Sound frequency and intensity.
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Structure and Function of Human Ear
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Sensing Body Position
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Lateral Line System
Sense objects that reflect pressure waves
and low frequency vibrations.
- Sensory structures within longitudinal
canal in fish’s skin that extends along
each side of the body and within several
canals in the head.
Sonar
Echolocation
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Lateral Line System
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Sensing Light: Vision
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Evolution of the Eye
Annelids, mollusks, arthropods, and
vertebrates have evolved well-developed,
image-forming eyes.
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Sensing Light: Vision
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Structure of the Eye
Cornea - Focuses light on covering
Lens - Completes focusing
Ciliary Muscles - Changes shape of lens
Iris - Regulates amount of light
Pupil - Transparent zone
Retina - Light sensitive receptor cells
Rods and Cones - Photoreceptors
- Fovea - Center of retina
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Human Eye
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Sensing Light: Vision
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Pigments in rods and cones are made from
carotenoids.
When it receives photon of light, pigment
undergoes change in shape.
- Must alter shape of attached opsin
protein.
Cis-retinal pigment absorbs light,
rotates sharply upward, forming transretinal form of pigment.
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Sensing Light: Vision
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Color Vision
Each of three kinds of cone cells possess
a different version of the opsin protein.
- Differences in shape affect flexibility of
attached retinal pigment, shifting
wavelength at which it absorbs light.
455 nanometers - blue
530 nanometers - green
625 nanometers - red
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Sensing Light: Vision
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Conveying Light Information to the Brain
Action potentials propagated along ganglion
cells are relayed through lateral geniculate
nuclei of thalamus and projected to occipital
lobe of the cerebral cortex.
- Pattern of activity among ganglion cells
across retina encodes a point-to-point
map of receptive field, allowing retina and
brain to image objects in visual space.
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Sensing Light: Vision
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Binocular Vision
When both eyes are trained on an object,
each eye views the object from a slightly
different view.
- Slight displacement provides ability to
perceive three-dimensional images and
sense depth.
Prey generally have eyes at side of
head, preventing binocular vision, but
enlarging overall perceptive fields.
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Binocular Vision
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Other Types of Sensory Reception
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Heat
Electricity
Magnetism
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Review
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Invertebrate Nervous System
Neurons
Nerve Impulses
Synapse
Drug Addiction
Evolution of Vertebrate Brain
Vertebrate Nervous System
Sensory Receptors
Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies
Copyright © McGraw-Hill Companies
Permission required for reproduction or display
Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies