Chapter 22 Notes
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Chapter 22
Descent with Modification:
A Darwinian View of Life
Question?
How
did the diversity of life
originate?
Through the process of
Evolution.
Evolution
The
processes that have
transformed life on earth
from its beginnings to today's
diversity.
“Nothing
in Biology Makes Sense
Except in the Light of Evolution”Theodosius Dobzhansky
Note:
Evolution does NOT
say anything about how life
originated
It DOES describe how life
changes through time.
Early Ideas on Life’s
Diversity
1. Plato - Organisms are
already perfectly adapted to
their environments.
2. Aristotle - Organisms
arranged on a “scale of life”
from simple to complex.
Result
No
evolution.
Life is already perfect and
doesn’t need to change.
All the rungs on life's "ladder"
are already occupied.
Carolus Linnaeus
(1707-1778)
Classified life according to genus and
species.
Thought the diversity in life was fixed from
life’s origins
Each lifeform was perfectly designed from
the beginning for its role
Still use his classification system today
Catastrophism
Georges
Cuvier
(1769-1832).
Attempted to
relate fossils to
current life.
Theory
Fossils
were the remains of
species lost due to
catastrophe.
No new species originated;
species could only be lost
over time.
Result - No evolution.
James Hutton
1795
- Gradualism
Profound change is the
cumulative product of slow,
but continuous processes.
Result
Changes
on the earth were
gradual, not catastrophic.
Charles Lyell
1797
- 1875.
Incorporated
Hutton’s
gradualism into a
theory called
Uniformitarianism.
Uniformitarianism
Geological
processes have
operated at the same rate
over the Earth’s history.
The lower the stratum, the more dissimilar life is to today’s life
Result
The
Earth must be VERY old.
(much older than 6000 years
of the fixed species concept).
Idea that slow and subtle
processes can cause
substantial change.
Jean Baptiste Lamarck
Published
theory in 1809.
Theory - Life
changed from
simple to
complex over
time.
Lamark
Fossils
were the remains of
past life forms.
Evolution did occur.
First to propose a
mechanism for evolution
Mechanisms
1. Use and Disuse Body
parts used to survive
become larger and stronger.
Body parts not used to survive
deteriorate.
Mechanisms
2. Acquired Characteristics
Modifications
acquired by
use/disuse were passed on to
offspring.
Problem
No
knowledge of genetics.
Acquired traits are not
transmitted offspring.
Lamarck’s Credits
Did
suggest correctly the role
of fossils in evolution.
Did suggest that adaptation
to the environment is a
primary product of evolution.
Thomas Malthus
Essay
on human population
growth in 1798.
Disease, famine, homelessness,
and war are inescapable because
human populations grow faster
than food supplies.
Darwin read Malthus.
Charles Darwin
Father
of the
modern theory
of evolution.
Theory Descent with
Modification.
Darwin's Background
Trained
as a Naturalist (after
trying religion and medicine).
Voyage of the Beagle
Result
Darwin's
training and travel
opportunities allowed him to
formulate and support his
ideas on Natural Selection.
On
twitter: @cdarwin
Darwin - 1859
Publication
of
"The Origin of
Species”
Comment
Darwin
best remembered for
the theory because of his
overwhelming evidence and
because he published.
Darwinian View
History
of life is like a tree
with branches over time from
a common source.
Current diversity of life is
caused by the forks from
common ancestors.
Example
“The Origin of Species”
Documented
the occurrence
of evolution.
Suggested that the
mechanism for evolution was
Natural Selection.
Observations:
Observation 1 – Members of a
population often vary greatly in
their traits.
.
Observation 2
Traits
are inherited from
parents to offspring.
Observation 3
All species are
capable of
producing
more offspring
than their
environment
can support.
Observation 4
Owing
to lack of food or
other resources, many of
these offspring do not
survive.
Inference 1
Individuals
whose inherited
traits give them a higher
probability of surviving and
reproducing in a given
environment tend to leave
more offspring than other
individuals.
Inference 2
This
unequal ability of
individuals to survive and
reproduce will lead to the
accumulation of favorable
traits in the population over
generations.
Nature
Determines
which
characteristics are favorable.
Determines who survives.
Result - “Natural Selection”
Natural Selection
in action
Artificial Selection
When
man determines the
characteristics that survive and
reproduce.
Result - the various breeds of
animals and plants we’ve
developed.
https://www.youtube.com/watch
?v=0jFGNQScRNY
Evolution Success
Measured By
Survival
Reproduction
Whoever
lives long enough
and has kids is the “winner”
in evolution.
Subtleties of Evolution
1. Populations are the units of
Evolution.
2. Only inherited characteristics
can evolve.
Comment
Acquired
characteristics may
allow a species to evolve
"outside" of Natural Selection.
Ex: culture, learning
Evidences for Evolution
Direct
observation of
evolutionary changes.
Fossils
Homology
Convergent Evolution
Biogeography
Molecular
Direct Observations
Drug
resistant HIV
Beak size in Birds
Many Others
HIV Drug Resistance
Drug
resistance strains
selected for by treatments
Result – resistant strains
became 100% dominant in 4-5
weeks.
Beak Size
Study – measured the
beak size of all birds in a
population over several
years.
Result – drought and food
competition changed beak
size.
Field
Fossils
Relics
or impressions of
organisms from the past.
Problem:
Show
changes over time from
simple to complex.
Many fossils don't have
descendants.
Evolution Viewpoint
Life
has
changed over
time.
Many species
failed to
survive and
became
extinct.
Comments
1. Fossilization is a rare event.
2. Only hard parts fossilize well.
3. Problem in finding fossils.
4. Interpretation.
5. Missing Links.
Homology
Homologysimilarities
resulting from
common
ancestors
Mammal forelimbs
Homology in Embryos
Problem
- closely related
organisms go through similar
stages in their embryonic
development.
Ex: Gill pouches in
vertebrates
Evolution Viewpoint
Ontogeny
is a replay of
Phylogeny.
(Development reflects descent)
Development reflects inheritance
of common control genes.
Problems
Vestigial Organs
- Rudimentary
structures of
marginal, if
any, use.
Whale Legs
Human Example
Evolution Viewpoint
Remodeling
of ancestral
structures as their functions
or adaptations changed.
Convergent Evolution
Unrelated
organisms show
similar adaptations.
Cause – lived in a similar
environment with similar
selection pressures.
Analogous Structures-similar function, but the structures are NOT from a
common ancestor
Biogeography
The
geographical distribution
of species.
Problem:
Species
mixtures on islands
Marsupials in Australia
Evolution Viewpoint
Biogeographical
patterns
reflect descent from the
ancestors that colonized that
area.
Molecular Biology
Study
of Evolution at the
DNA or protein levels.
Related
species have similar
DNA sequences.
Evolution Viewpoint
Related
species share a
common ancestrial DNA.
The closer the relationship,
the more similar the DNA
sequences should be.
Summary
Darwin's
ideas now a "Theory”.
Predictions of a Theory are
tested by experiments and
observations.
Be familiar with the pre-Darwin
views of evolution.
Summary
Know
Darwin’s “observations”
and “inferences”.
Be able to discuss the various
evidences of Darwinian
evolution.