Transcript CCEB
Pharmacogenetics of
Leukemia Treatment
Response
Richard Aplenc
May 2nd, 2008
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Pediatric Leukemia
Most common pediatric malignancy
Four types
ALL
AML
CML
JMML
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Leukemia Treatment
Varies both by disease and treating group
Generally curable
~80% in ALL
~60% in AML
Toxicity important
Long term effects in ALL
Infection and cardiac toxicity in AML
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Leukemia Treatment
Multi-agent
Over time
Substantial impact on patient and family
Accurate response prediction is clinically
very important
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ALL Therapy
Induction
L-Asp
Consolidation
Steroids
MTX
Interim
Maintenance
VCR
6-MP/6-TG
Delayed
Intensification
AraC
Maintenance
Doxorubicin Cyclophosphamide
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Predicting Treatment Response
Leukemic blast characteristics
Morphology
Cytogenetics
Molecular alterations (BCR-ABL)
Patient characteristics
Age
Gender
Genetic information?
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Genetic Information
Variation in DNA sequence throughout the
genome
Types of variation include
Gene deletions (GSTT1)
Duplications of DNA regions (TS 28 bp)
Changes in single base pairs (SNPs)
Allele, genotype, haplotype
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Allele/Genotype/Haplotype/CNV
SNP: Single Nucleotide Polymorphism
An allele is a single value for a single
marker
A genotype is a pair of alleles for a given
marker and both chromosomes in a
single person
Copy number variation (CNV) of DNA
sequences
Genotype
SNP 1
SNP 2
Haplotype
A haplotype is an ordered series of
alleles for many markers on a single
chromosome
Allele
...
Chromosome Chromosome from
from one parent
other parent
SNP example:
G
GTACGTTCG GGGCGGGAT
T
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Impact of Genetic Variability
Loss of gene = loss of function
Duplication of DNA segments and single
base pair changes may have different
effects depending on position
Gain of function, loss of function, no change
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Our Dream
One Genotype Would Explain
Treatment Response
Why Did We Have This Dream?
Thiopurine methylatransferase (TPMT)
Low frequency variants have complete loss of
thiopurine metabolizing abilities
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That Dream Has Ended
Why Is That?
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TPMT
One Gene, One Pathway, One Exposure
SH
TX
TU
HO
Allopurinol
N
N
SH
OH
N
N
SH
N
N
TPMT
SCH 3
N
N
N
HO
HGPRT
N
N
PO 4 CH 2
TIMP
TXMP
TGMP
O
H
N
N
N
XO
H
N
N
Mercaptopurine
HO
OH
TPMT Deficiency
H
6-MMP
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Two Remaining Questions
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Question 1:
Can we utilize data on host
genetic variability in a clinically
meaningful way?
Question 2:
Is Theo Zaoutis really Neo?
This Makes Sense Because…
Lisa Z looks like
Trinity
And Because…
Paul Offit is clearly
Morpheus
Now That Everyone is Awake…
Return to Question 1
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Moving Towards the Answer
Decide on the question
Understand the complex phenotype issues
Host genetics
Environment
Address the genetic epidemiology issues
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What is the Question?
Does the genotype inform us of the
biology underlying a clinical outcome?
Etiology
Does the genotype predict a clinical
outcome?
Prediction
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One Conceptual Approach
Etiology
Sensitivity
Probability of positive test given disease
Prediction
Positive predictive value
Probability of disease given positive test
Seems obvious but impacts analysis
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Complex Phenotype: Host Genetics
Common SNPs will have modest effects
Potentially large impact for the population
Rare SNPs may have bigger effects
Small population impact
SNP frequency and the effect size
determine sample size
SNP frequency varies by ethnicity
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Complex Phenotype: Environment
Identify and measure relevant covariates
Genotype does not matter if the patient
doesn’t take the medication
Concomitant medications
Drug-drug interactions
Alternative medications
Folic acid supplimentation
Other environmental exposures
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What are the Genetic Epidemiology
Issues?
Population stratification
Variation of SNP frequency by ethnicity
High dimensional data
Gene-environment interactions
Interaction of host genetics with environment
Gene-gene interactions
Interaction of different SNPs
Multiple comparisons
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Some Examples from Our Data
Methotrexate interrupts the folate cycle
ALL blasts are sensitive to folate depletion
Polymorphisms in genes in the folate cycle
may impact methotrexate efficacy
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Relapse Free Survival by MTHFR C677T Variant Allele
1.00
0.90
0.80
0.70
0.60
p = 0.0486
0.50
0
5
10
Years
Wildtype (C)
Variant (T)
MTHFR C677T Cox Model
Covariate
HR
p
95% CI
C677T variant
1.93
0.004
1.229
3.037
Day 7 BM
1.77
0.013
1.125
2.773
Age
1.11
0.016
1.020
1.220
Race
1.71
0.307
0.610
4.798
Gender
1.37
0.238
0.811
2.323
Rx Arm
1.18
0.214
0.908
1.535
WBC
0.99
0.335
0.971
1.010
Phenotype
0.95
0.776
0.661
1.362
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MTHFR C677T and Infection Risk
Gene
MTHFR
C677T
MTHFR
C677T
MTHFR
C677T
Genotype
N
C/C
C/T
T/T
C/C
C/T
T/T
C/C
C/T
T/T
224
187
72
224
187
72
224
187
72
Num.
Infection Type
Infection
46
Sepsis
42
Sepsis
16
Sepsis
155
Fever/Neutropenia
120
Fever/Neutropenia
53
Fever/Neutropenia
123
Infection - Other
113
Infection - Other
43
Infection - Other
OR
1
1.13
1.13
1
0.83
1.32
1
1.27
1.2
95% CI
P value
0.700-1.818
0.585-2.188
0.86
0.546-1.276
0.709-2.447
0.34
0.850-1.887
0.690-2.087
0.49
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MTHFR Conclusions
The MTHFR C677T variant allele seems to
impact relapse risk
Dose adjustment of methotrexate for
toxicity/infection does not ameliorate this effect
Dose adjustment based on genotype may be
clinically useful
Replication in anther sample set is ongoing
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MTFHR Issues
Allele versus genotype versus haplotype
Clinically meaningful analysis
Positive predictive value
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Relapse Free Survival by MTHFR C677T Variant Allele
1.00
0.90
0.80
0.70
0.60
p = 0.0486
0.50
0
5
10
Years
Wildtype (C)
Variant (T)
Relapse Free Survival by MTHFR C677T Genotype
1.00
0.90
0.80
0.70
0.60
CC vs TT, p = 0.0477
0.50
0
5
10
Years
Wildtype (CC)
Variant (TT)
Heterozygote (CT)
Kaplan-Meier survival estimates, by haplo
1.00
0.75
0.50
0.25
0.00
0
5
10
analysis time
CA CA
CG CG
TA CG
TA TG
TG TG
CA CG
TA CA
TA TA
TG CG
15
PPV with Time to Relapse Data
This is the metric of interest to oncologists
Moscowitz and Pepe defined positive
predictive value in survival time data
PPVXk(t) = P(T ≤ t | Xk = 1)
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PPV Conclusions
Although statistically significant, the
MTHFR C677T allele has a PPV of 35%
This is worse than flipping a coin
Important question is the increased predictive
value above baseline
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TS 28 bp as Example
N
RFS
HR
CI
p
2R/2R
83
80%
1
--
--
2R/3R
196
79%
1.68
0.863-3.255
0.13
3R/3R
103
73%
1.87
0.942-3.721
0.074
3R/4R
20
60%
3.69
1.436-9.481
0.007
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TS 28 bp Bootstrapping
Does knowledge of TS genotype improve
prediction of relapse?
Bootstrap comparison of relapse free
survival of all patients with those with
particular TS polymorphisms
No additional predictive value from
knowing TS genotype
Caveat of sample size issues
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Other Genetic Epidemiology
Issues
Multiple comparisons
Gene-gene and gene-environment
interactions
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Multiple Comparisons
Probability of finding a false association by
chance = 1 - 0.95n
n = 10, p = 40%
n = 100, p = 99.4%
Our data:
19 genotypes, 2 genders, 3 different relapse
sites
N = 228, p = 99.99959%
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Methods for Multiple Comparisons
Ignore it
Validation sample set
Adjust p-values
Bonferroni
False discovery rate (FDR) Benjamini et al 2001
Use Bayesian methods
False positive report probability (FPRP) Wacholder et al 2004
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High Dimensional Data
The number of cells (N) needed to split R
variables into X partitions:
R
N=X
A single 2-way combination
R = 2, X= 3, N= 9
We have evaluated 19 genotypes
All 2-way combinations of our genotypes
R = 19, X = 3, N = 1,162,261,467
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High Dimensional Data Methods
Several methods in current use
We have used patterning with recursive
partitioning (CART)
Create groups as uniform as possible
Use with genotype and other covariates
No p-values
Confirmation by cross-validation within the
sample set
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CART Caveats
No p-values
Need to validate in a separate sample
Often difficult to interpret results,
particularly of higher order interactions
i.e. 2 genotypes and 1 environmental factor
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Future Directions
Validate and extend genotyping in another
ALL sample set
Incorporate drug dose data
Investigate the impact of genetic variability
on infection risk in pediatric myeloid
leukemia
R01 resubmission with Theo Zaoutis
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The End….
Thanks to everyone who makes it safe to swim with the sharks. Bev Lange,
Tim Rebbeck,Jinbo Chen, Theo Zaoutis, Tom McWilliams, Peggy Han,
Shannon Smith, Michelle Horn, Melanie Doran. Funded by RO1 CA108862-01.
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